Samantha Markle furiously blasted sister Meghan's "ludicrous" law suit and warned that the "truth" would come out.
The 55-year-old appeared on This Morning today via video link in a bid to defend dad Thomas amid the news that her father would testify against sister Meghan in court.
She also demanded that Meghan and Harry apologise to Thomas and accused them of setting up a "smear campaign" against him.
Samantha also confirmed that her father would be willing to testify in court against Meghan.
In October, The Duchess of Sussex took legal action against the Mail on Sunday over its decision to publish a highly personal letter which she penned for her dad.
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But the estranged father and daughter appear to be on a collision course after reports that the newspaper signed up Meghan’s father as its star witness.
Speaking from Tampa, Florida, Samantha was asked by hosts Eamonn Holmes and Ruth Langsford whether Thomas Markle would be willing to testify.
"Well of course he would," she said. "The truth is the truth. Based on that, of course he would testify
firstly she had mailed him the letter… in the US if someone mails a letter and it lands in your postal box it's your property."
Pointing out that there was the "issue that she shared it with several of her friends" and that "it went to the public forum", she added: "This claim of privacy doesn't seem to have a legitimate basis. It seems rather ludicrous."
The Mail on Sunday will argue there is a huge public interest in the personal relationships of members of the royal family as part of its defence, court documents show.
Meghan's lawyers filed a High Court claim against the paper and its parent company Associated Newspapers over the alleged misuse of private information.
But in its legal defence the paper claims the duchess "did not have a reasonable expectation of privacy that the contents of the letter were private and would remain so", according to the BBC.
*This Morning airs weekdays at 10am on ITV
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